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Structures of monomeric and oligomeric forms of the Toxoplasma gondii perforin-like protein 1

Abstract:
Copyright © 2018 The Authors, some rights reserved. Toxoplasma and Plasmodium are the parasitic agents of toxoplasmosis and malaria, respectively, and use perforinlike proteins (PLPs) to invade host organisms and complete their life cycles. The Toxoplasma gondii PLP1 (TgPLP1) is required for efficient exit from parasitophorous vacuoles in which proliferation occurs. We report structures of the membrane attack complex/perforin (MACPF) and Apicomplexan PLP C-terminal b-pleated sheet (APCb) domains of TgPLP1. The MACPF domain forms hexameric assemblies, with ring and helix geometries, and the APCb domain has a novel b-prism fold joined to the MACPF domain by a short linker. Molecular dynamics simulations suggest that the helical MACPF oligomer preserves a biologically important interface, whereas the APCb domain binds preferentially through a hydrophobic loop to membrane phosphatidylethanolamine, enhanced by the additional presence of inositol phosphate lipids. This mode of membrane binding is supported by site-directed mutagenesis data from a liposome-based assay. Together, these structural and biophysical findings provide insights into the molecular mechanism of membrane targeting by TgPLP1.
Publication status:
Published
Peer review status:
Peer reviewed

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Publisher copy:
10.1126/sciadv.aaq0762

Authors


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Institution:
University of Oxford
Division:
Medical Sciences Division
Department:
NDM; Structural Biology
Role:
Author
More by this author
Institution:
University of Oxford
Division:
Medical Sciences Division
Department:
NDM; Structural Biology
Role:
Author
More by this author
Institution:
University of Oxford
Division:
Medical Sciences Division
Department:
NDM; Structural Biology
Role:
Author


More from this funder
Funding agency for:
Rezelj, S
Anderluh, G
Grant:
P1-0391
P1-0391
More from this funder
Funding agency for:
Williams, S
Grant:
BB/M011224/1
BB/R002517/1
More from this funder
Funding agency for:
Williams, S
Grant:
BB/M011224/1


Publisher:
American Association for the Advancement of Science
Journal:
Science Advances More from this journal
Volume:
4
Issue:
3
Article number:
eaaq0762
Publication date:
2018-03-21
Acceptance date:
2018-02-09
DOI:
ISSN:
2375-2548


Keywords:
Pubs id:
pubs:833617
UUID:
uuid:9499dc1f-82ec-4bb6-badf-27ab8d6ac7e4
Local pid:
pubs:833617
Source identifiers:
833617
Deposit date:
2018-04-08

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