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Non‐equivalence in old‐ and new‐flagellum daughter cells of a proliferative division in Trypanosoma brucei

Abstract:
Differentiation of Trypanosoma brucei, a flagellated protozoan parasite, between life cycle stages typically occurs through an asymmetric cell division process, producing two morphologically distinct daughter cells. Conversely, proliferative cell divisions produce two daughter cells, which look similar but are not identical. To examine in detail differences between the daughter cells of a proliferative division of procyclic T. brucei we used the recently identified constituents of the flagella connector. These segregate asymmetrically during cytokinesis allowing the new‐flagellum and the old‐flagellum daughters to be distinguished. We discovered that there are distinct morphological differences between the two daughters, with the new‐flagellum daughter in particular re‐modelling rapidly and extensively in early G1. This re‐modelling process involves an increase in cell body, flagellum and flagellum attachment zone length and is accompanied by architectural changes to the anterior cell end. The old‐flagellum daughter undergoes a different G1 re‐modelling, however, despite this there was no difference in G1 duration of their respective cell cycles. This work demonstrates that the two daughters of a proliferative division of T. brucei are non‐equivalent and enables more refined morphological analysis of mutant phenotypes. We suggest all proliferative divisions in T. brucei and related organisms will involve non‐equivalence.
Publication status:
Published
Peer review status:
Peer reviewed

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Publisher copy:
10.1111/mmi.14345

Authors


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Institution:
University of Oxford
Oxford college:
Lincoln College
Role:
Author
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Institution:
University of Oxford
Department:
Pathology Dunn School
Role:
Author
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Institution:
University of Oxford
Division:
MSD
Department:
Pathology Dunn School
Role:
Author
ORCID:
0000-0002-4270-8360


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Grant:
211075/Z/18/Z
103261/Z/13/Z
104627/Z/14/Z


Publisher:
Wiley
Journal:
Molecular Microbiology More from this journal
Volume:
112
Issue:
3
Pages:
1024-1040
Publication date:
2019-07-25
Acceptance date:
2019-07-08
DOI:
EISSN:
1365-2958
ISSN:
0950-382X
Pmid:
31286583


Language:
English
Keywords:
Pubs id:
pubs:1032322
UUID:
uuid:f37176f2-9ad3-4b6a-9dda-5ea18859dc02
Local pid:
pubs:1032322
Source identifiers:
1032322
Deposit date:
2019-07-26

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