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Journal article

Local unfolding of the HSP27 monomer regulates chaperone activity

Abstract:
The small heat-shock protein HSP27 is a redox-sensitive molecular chaperone that is expressed throughout the human body. Here we describe redox-induced changes to the structure, dynamics, and function of HSP27 and its conserved α-crystallin domain (ACD), and provide the first structural characterization of a small heat-shock protein monomer using nuclear magnetic resonance (NMR) spectroscopy. While HSP27 assembles into oligomers, we show that the transiently populated monomers that are released upon reduction are highly active chaperones in vitro, but are kinetically unstable and susceptible to uncontrolled self-aggregation. By using relaxation dispersion and high-pressure NMR spectroscopy, we observe that the pair of β-strands that mediate dimerization partially unfold in the monomer. Strikingly, we note that numerous HSP27 mutations associated with inherited neuropathies cluster to this dynamic region. The high degree of sequence conservation in the ACD amongst mammalian sHSPs suggests that the exposed, disordered interface within the context of oligomers or free in solution may be a general, functional feature of these molecular chaperones.
Publication status:
Published
Peer review status:
Peer reviewed

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Publisher copy:
10.1038/s41467-019-08557-8

Authors


More by this author
Institution:
University of Oxford
Division:
MPLS Division
Department:
Chemistry
Sub department:
Physical & Theoretical Chem
Role:
Author
More by this author
Institution:
University of Oxford
Division:
MPLS Division
Department:
Chemistry
Sub department:
Physical & Theoretical Chem
Role:
Author
More by this author
Institution:
University of Oxford
Division:
MPLS Division
Department:
Chemistry
Sub department:
Physical & Theoretical Chem
Role:
Author
More by this author
Institution:
University of Oxford
Division:
MPLS Division
Department:
Chemistry
Sub department:
Physical & Theoretical Chem
Role:
Author


Publisher:
Springer Nature
Journal:
Nature Communications More from this journal
Volume:
10
Article number:
1068
Publication date:
2019-03-06
Acceptance date:
2019-01-05
DOI:
EISSN:
2041-1723


Keywords:
Pubs id:
pubs:965607
UUID:
uuid:d2a68ded-19e8-4498-97a2-85c01d542ed1
Local pid:
pubs:965607
Source identifiers:
965607
Deposit date:
2019-01-22

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