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The evolution of short- and long-range weapons for bacterial competition

Abstract:
Bacteria possess a diverse range of mechanisms for inhibiting competitors, including bacteriocins, tailocins, type VI secretion systems and contact-dependent inhibition (CDI). Why bacteria have evolved such a wide array of weapon systems remains a mystery. Here we develop an agent-based model to compare short-range weapons that require cell-cell contact, with long-range weapons that rely on diffusion. Our model predicts that contact weapons are useful when an attacking strain is outnumbered, facilitating invasion and establishment. By contrast, ranged weapons tend to be effective only when attackers are abundant. We test our predictions with the opportunistic pathogen Pseudomonas aeruginosa, which naturally carries multiple weapons, including CDI and diffusing tailocins. As predicted, short-range CDI can function at low and high frequencies, while long-range tailocins require high frequency and cell density to function effectively. Head-to-head competition experiments with the two weapon types further support our predictions: a tailocin attacker defeats CDI only when it is numerically dominant, but then we find it can be devastating. Finally, we show that the two weapons work well together when one strain employs both. We conclude that short- and long-range weapons serve different functions and allow bacteria to fight both as individuals and as a group.
Publication status:
Published
Peer review status:
Peer reviewed

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Publisher copy:
10.1038/s41559-023-02234-2

Authors


More by this author
Institution:
University of Oxford
Division:
MPLS
Department:
Biology
Role:
Author
ORCID:
0000-0003-2552-5324
More by this author
Role:
Author
ORCID:
0000-0002-9270-7614
More by this author
Institution:
University of Oxford
Division:
MPLS
Department:
Biology
Oxford college:
Magdalen College
Role:
Author
ORCID:
0000-0003-4687-6633


Publisher:
Springer Nature
Journal:
Nature Ecology and Evolution More from this journal
Volume:
7
Issue:
12
Pages:
2080–2091
Place of publication:
England
Publication date:
2023-11-30
Acceptance date:
2023-09-22
DOI:
EISSN:
2397-334X
ISSN:
2397-334X
Pmid:
38036633


Language:
English
Keywords:
Pubs id:
1574538
Local pid:
pubs:1574538
Deposit date:
2023-12-12

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