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Evolution of a plant growth-regulatory protein interaction specificity

Abstract:

Specific protein–protein interactions (PPIs) enable biological regulation. However, the evolution of PPI specificity is little understood. Here we trace the evolution of the land-plant growth-regulatory DELLA–SLY1/GID2 PPI, revealing progressive increase in specificity of affinity of SLY1/GID2 for a particular DELLA form. While early-diverging SLY1s display relatively broad-range DELLA affinity, later-diverging SLY1s tend towards increasingly stringent affinity for a specific DELLA A’ form generated by the growth-promoting phytohormone gibberellin (GA). Our novel mutational strategy reveals amino acid substitutions contributing to the evolution of Arabidopsis thaliana SLY1 A’ specificity, also showing that routes permitting reversion to broader affinity became increasingly constrained over evolutionary time. We suggest that progressive affinity narrowing may be an important evolutionary driver of PPI specificity and that increase in SLY1/GID2-DELLA specificity enabled the enhanced flexibility of plant physiological environmental adaptation conferred by the GA-DELLA growth-regulatory mechanism.

Publication status:
Published
Peer review status:
Peer reviewed

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Publisher copy:
10.1038/s41477-023-01556-0

Authors

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Institution:
University of Oxford
Division:
MPLS
Department:
Biology
Role:
Author
ORCID:
0000-0002-0651-2598
More by this author
Institution:
University of Oxford
Division:
MPLS
Department:
Biology
Oxford college:
Wolfson College
Role:
Author
ORCID:
0000-0003-0892-100X
More by this author
Role:
Author
ORCID:
0000-0002-6943-5171
More by this author
Institution:
University of Oxford
Division:
MPLS
Department:
Biology
Role:
Author
ORCID:
0000-0002-6617-0112
More by this author
Role:
Author
ORCID:
0000-0003-2837-9027


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Funder identifier:
https://ror.org/01h0zpd94
Grant:
32122065
More from this funder
Funder identifier:
https://ror.org/00cwqg982
Grant:
BB/S013741/1


Publisher:
Springer Nature
Journal:
Nature Plants More from this journal
Volume:
9
Issue:
12
Pages:
2059-2070
Publication date:
2023-10-30
Acceptance date:
2023-09-27
DOI:
EISSN:
2055-0278
ISSN:
2055-026X
Pmid:
37903985


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