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Intraspecific variation in symbiont density in an insect-microbe symbiosis

Abstract:
Many insects host vertically transmitted microbes, which can confer benefits to their hosts but are costly to maintain and regulate. A key feature of these symbioses is variation: for example, symbiont density can vary among host and symbiont genotypes. However, the evolutionary forces maintaining this variation remain unclear. We studied variation in symbiont density using the pea aphid (Acyrthosiphon pisum) and the bacterium Regiella insecticola, a symbiont that can protect its host against fungal pathogens. We found that relative symbiont density varies both between two Regiella phylogenetic clades and among aphid “biotypes.” Higher density symbiont infections are correlated with stronger survival costs, but variation in density has little effect on the protection Regiella provides against fungi. Instead, we found that in some aphid genotypes, a dramatic decline in symbiont density precedes the loss of a symbiont infection. Together, our data suggest that the optimal density of a symbiont infection is likely different from the perspective of aphid and microbial fitness. Regiella might prevent loss by maintaining high within‐host densities, but hosts do not appear to benefit from higher symbiont numbers and may be advantaged by losing costly symbionts in certain environments. The standing variation in symbiont density observed in natural populations could therefore be maintained by antagonistic coevolutionary interactions between hosts and their symbiotic microbes.
Publication status:
Published
Peer review status:
Peer reviewed

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Publisher copy:
10.1111/mec.15821

Authors


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Role:
Author
ORCID:
0000-0002-0679-4732
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Institution:
University of Oxford
Division:
SSD
Role:
Author


Publisher:
Wiley
Journal:
Molecular Ecology More from this journal
Volume:
30
Issue:
6
Pages:
1559-1569
Publication date:
2021-02-11
Acceptance date:
2021-01-20
DOI:
EISSN:
1365-294X
ISSN:
0962-1083
Pmid:
33512733


Language:
English
Keywords:
Pubs id:
1159363
Local pid:
pubs:1159363
Deposit date:
2021-02-22

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